Epistasis 1 Epistasis, Genetic 1 Gene Frequency 1 Genetic Variation 1 Genetic variances Inbreeding 1 Linkage Disequilibrium 1 Linkage disequilibrium 1 Models, Genetic 1 additive effect 1 article 1 biological model 1 covariance 1 dominant gene 1 epistasis 1 gene frequency 1 gene linkage disequilibrium 1 genetic variability 1 genetic variation 1 inbreeding 1 simulation 1